{"id":52954,"date":"2025-08-17T19:03:41","date_gmt":"2025-08-17T10:03:41","guid":{"rendered":"https:\/\/ichihashi.me\/wp\/?p=52954"},"modified":"2026-04-08T08:13:47","modified_gmt":"2026-04-07T23:13:47","slug":"%e3%82%a4%e3%83%8d%e7%9b%aepoales%e3%81%ae%e7%b3%bb%e7%b5%b1%e6%a6%82%e8%a6%81","status":"publish","type":"post","link":"https:\/\/ichihashi.me\/wp\/?p=52954","title":{"rendered":"<span id=\"Top\">\u30a4\u30cd\u76ee(Poales)<\/span>\u306e\u7cfb\u7d71\u6982\u8981"},"content":{"rendered":"<p>\u3000\u30a4\u30cd\u76ee(Poales)\u306f\uff12\u4e07\u7a2e\u4ee5\u4e0a\u304c\u8a18\u8f09\u3055\u308c\u3066\u6765\u305f\u5927\u898f\u6a21\u306a\u30b0\u30eb\u30fc\u30d7\u3067\u3001\u305d\u306e\u3046\u30611\u4e072000\u7a2e\u4ee5\u4e0a\u304c\u30a4\u30cd\u79d1\u306e\u6240\u5c5e\u3068\u306a\u3063\u3066\u3044\u307e\u3059(Wu <i>et al.<\/i>,2022)\u3002\u30a4\u30cd\u76ee\u5185\u306e\u30ab\u30c4\u30de\u30bf\u30bd\u30a6\u5468\u8fba\u306b\u8a8d\u3081\u3089\u308c\u305f3\u79d1\u304c\u7d71\u5408\u3055\u308c\u305f\u7d50\u679c\u3001APG-IV(2016)\u3067\u306f14\u306e\u79d1\u306b\u6574\u7406\u3055\u308c\u3066\u3044\u307e\u3059\u3002<br \/>\n\u3000\u30a4\u30cd\u79d1(Poaceae)\u306e\u5206\u985e\u306b\u95a2\u3057\u3066\u306f\u3001<a href=\"#GPWG_2001\">Grass Phtlogeny Working Group(2001)<\/a>\u306e\u516c\u8868\u304b\u3089\u3082\u3046\u3059\u3050\u56db\u534a\u4e16\u7d00\u3068\u306a\u308a\u3001\u79d1\u5185\u306e\u3075\u305f\u3064\u306e\u5206\u5c90\u7fa4(PACMAD\u5206\u5c90\u7fa4\u3001BEP\u5206\u5c90\u7fa4)\u3001\u7279\u306b\u5f8c\u8005\u304c\u5358\u7cfb\u3067\u3042\u308b\u3053\u3068\u306f\u5e83\u304f\u53d7\u3051\u5165\u308c\u3089\u308c\u3066\u3044\u307e\u3059\u304c\u3001\u9023\u306e\u30ec\u30d9\u30eb\u4ee5\u4e0b\u306b\u306f\u307e\u3060\u89e3\u6c7a\u3059\u3079\u304d\u5206\u5c90\u304c\u591a\u6570\u6b8b\u3055\u308c\u3066\u3044\u308b\u69d8\u3067\u3059\u3002<br \/>\n<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640\">\u30a4\u30cd\u79d1\u306e\u5199\u771f\u6574\u7406<\/a>\u3000<a href=\"https:\/\/ichihashi.me\/wp\/?p=34884\">\u30bf\u30b1\u4e9c\u79d1\u306e\u5199\u771f<\/a><br \/>\n<br \/>\n\u7dad\u7ba1\u675f\u690d\u7269\u306e\u5206\u985e\u3010<a href=\"https:\/\/ichihashi.me\/wp\/?p=46395\">\u6982\u8981<\/a>\u3001<a href=\"https:\/\/ichihashi.me\/wp\/?p=42398\">\u5c0f\u8449\u690d\u7269\u3068\u5927\u8449\u30b7\u30c0\u690d\u7269<\/a>\u3001<a href=\"https:\/\/ichihashi.me\/wp\/?p=42340#Neontology\">\u88f8\u5b50\u690d\u7269<\/a>\u3001<a href=\"https:\/\/ichihashi.me\/wp\/?p=40497\">\u88ab\u5b50\u690d\u7269(APG-IV\u4f53\u7cfb)<\/a>\u3011\u3010<a href=\"https:\/\/ichihashi.me\/wp\/?p=44648\">\u751f\u7269\u7cfb\u7d71\u6a39<\/a>\u3011<\/p>\n<hr id=\"Poales_Phylogeny\">\n\u3010\u30a4\u30cd\u76ee(Poales)\u306e\u7cfb\u7d71\u6982\u8981\u3011<a href=\"#Wu_2022\"> Wu <i>et al.<\/i>(2022)<\/a><br \/>\n<span style=\"color:green; background-color:inherit; font-size:small;\">\u3000<a href=\"#APG-IV\">APG-IV(2016)<\/a>\u3067\u306f\u3001\u30ab\u30c4\u30de\u30c0\u30bd\u30a6\u79d1\u3068\u30a2\u30ca\u30eb\u30c8\u30ea\u30a2\u79d1\u306f\u30b5\u30f3\u30a2\u30bd\u30a6\u79d1\u306b\u7d71\u5408\u3055\u308c\u3001\u30a4\u30cd\u76ee\u306f14\u79d1\u3068\u306a\u3063\u3066\u3044\u307e\u3059\u3002<\/span><\/p>\n<div style=\"line-height: 1em; font-family: monospace; padding:0 0 1em 0;\">\n\u3000\u3000\u3000\u3000\u3000\u3000\u250c<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30b0\u30e9\u30df\u30cb\u30c9\u5206\u5c90\u7fa4(Graminids)<\/span><br \/>\n\u3000\u3000\u3000\u3000\u3000\u3000\u2502\u2502\u3000\u250c<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640\">\u30a4\u30cd\u79d1(Poaceae)<\/a> \u2500\u2192 \u3010<a href=\"#Poaceae_Phylogeny_GPWG-II\">\u30a4\u30cd\u79d1<\/a>\u3011<br \/>\n\u3000\u3000\u3000\u3000\u3000\u250c\u2524\u2502\u250c\u2534\u252c\u30b8\u30e7\u30a4\u30f3\u30d3\u30ec\u30a2\u79d1(Joinvilleaceae)<br \/>\n\u3000\u3000\u3000\u3000\u3000\u2502\u2502\u2514\u2524\u3000\u2514\u30a8\u30af\u30c7\u30a4\u30aa\u30b3\u30ec\u30a2\u79d1(Ecdeiocoleaceae)<br \/>\n\u3000\u3000\u3000\u3000\u250c\u2524\u2502\u3000\u2514\u30c8\u30a6\u30c4\u30eb\u30e2\u30c9\u30ad\u79d1(Flagellariaceae)<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u2514<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30b5\u30f3\u30a2\u30bd\u30a6\u5206\u5c90\u7fa4(Restiids)<\/span> = \u73fe\u30b5\u30f3\u30a2\u30bd\u30a6\u79d1<br \/>\n\u3000\u3000\u3000\u250c\u2524\u2502\u3000\u2502\u3000\u250c\u30ab\u30c4\u30de\u30c0\u30bd\u30a6(\u4e9c)\u79d1(Centrolepidaceae)<br \/>\n\u3000\u3000\u3000\u2502\u2502\u2502\u3000\u2514\u252c\u2534\u30b5\u30f3\u30a2\u30bd\u30a6\u79d1(Restionaceae)<br \/>\n\u3000\u3000\u250c\u2524\u2502\u2502\u3000\u3000\u2514\u30a2\u30ca\u30eb\u30c8\u30ea\u30a2(\u4e9c)\u79d1(Anarthriaceae)<br \/>\n\u3000\u3000\u2502\u2502\u2502\u2514\u30c8\u30a6\u30a8\u30f3\u30bd\u30a6\u79d1(Xyridaceae)\u3000\u2510<br \/>\n\u3000\u250c\u2524\u2502\u2514\u30db\u30b7\u30af\u30b5\u79d1(Eriocaulaceae)\u3000\u3000 \u251c<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30c8\u30a6\u30a8\u30f3\u30bd\u30a6\u6bb5\u968e\u7fa4<\/span><br \/>\n\u3000\u2502\u2502\u2514\u30de\u30e4\u30ab\u79d1(Mayacaceae)\u3000\u3000\u3000\u3000\u3000\u3000\u2518<span style=\"color:maroon; background-color:inherit; font-weight:700;\">(Xyrids)<\/span><br \/>\n\u3000\u2502\u2514<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30ab\u30e4\u30c4\u30ea\u30b0\u30b5\u5206\u5c90\u7fa4(Cyprids)<\/span><br \/>\n\u250c\u2524\u3000\u2502\u3000\u250c<a href=\"https:\/\/ichihashi.me\/wp\/?p=40631\">\u30ab\u30e4\u30c4\u30ea\u30b0\u30b5\u79d1(Cyperaceae)<\/a><br \/>\n\u2502\u2502\u3000\u2502\u250c\u2534<a href=\"https:\/\/ichihashi.me\/wp\/?p=40625\">\u30a4\u30b0\u30b5\u79d1(Juncaceae)<\/a><br \/>\n\u2524\u2502\u3000\u2514\u2534\u30c8\u30a5\u30eb\u30cb\u30a2\u79d1(Thurniaceae)<br \/>\n\u2502\u2514<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u57fa\u90e8\u30a4\u30cd\u76ee(Basal Poales)<\/span><br \/>\n\u2502\u3000\u2502\u3000\u250c\u30e9\u30d1\u30c6\u30a2\u79d1(Rapateaceae)<br \/>\n\u2502\u3000\u2502\u250c\u2534<a href=\"https:\/\/ichihashi.me\/wp\/?p=40621\">\u30d1\u30a4\u30ca\u30c3\u30d7\u30eb\u79d1(Bromeliaceae)<\/a><br \/>\n\u2502\u3000\u2514\u2534<a href=\"https:\/\/ichihashi.me\/wp\/?p=40618\">\u30ac\u30de\u79d1(Typhaceae)<\/a><br \/>\n\u2514<span style=\"font-weight:700;\">\u5916\u7fa4(Outgroup)<\/span><br \/>\n\u3000\u2514\u252c<a href=\"https:\/\/ichihashi.me\/wp\/?p=40497#Commelinids\">\u30c4\u30e6\u30af\u30b5\u76ee(Commelinales)<\/a><br \/>\n\u3000\u3000\u2514<a href=\"https:\/\/ichihashi.me\/wp\/?p=40497#Commelinids\">\u30b7\u30e7\u30a6\u30ac\u76ee(Zingiberales)<\/a>\n<\/div>\n<p><!--\n<a href=\"#Top\">To Top<\/a>\n\n\n<hr id=\"Poaceae_Phylogeny\">\n\u3010\u30a4\u30cd\u79d1(Poaceae)\u306e\u7cfb\u7d71\u6982\u8981\u3011<a href=\"# Kellogg_2009\"> Kellogg(2009)<\/a><br \/>\n\n\n<div style=\"line-height: 1em; font-family: monospace; padding:0 0 1em 0;\">\n\u3000\u3000\u250c\u30ab\u30c4\u30de\u30c0\u30bd\u30a6(\u4e9c)\u79d1(Centrolepidaceae)\u2510<br \/>\n\u250c\u252c\u2534\u30b5\u30f3\u30a2\u30bd\u30a6\u79d1(Restionaceae)\u3000\u3000\u3000\u3000\u3000\u251c\u73fe\u30b5\u30f3\u30a2\u30bd\u30a6\u79d1<br \/>\n\u2524\u2514\u30a2\u30ca\u30eb\u30c8\u30ea\u30a2(\u4e9c)\u79d1(Anarthriaceae)\u3000\u3000 \u2518(Restionaceae)<br \/>\n\u2502\u250c\u30c8\u30a6\u30c4\u30eb\u30e2\u30c9\u30ad\u79d1(Flagellariaceae)<br \/>\n\u2514\u2524\u250c\u252c\u30a8\u30af\u30c7\u30a4\u30aa\u30b3\u30ec\u30a2\u79d1(Ecdeiocoleaceae)<br \/>\n\u3000\u2514\u2524\u2514\u30b8\u30e7\u30a4\u30f3\u30d3\u30ec\u30a2\u79d1(Joinvilleaceae)<br \/>\n\u3000\u3000\u2514\u252c\u30a2\u30ce\u30e2\u30af\u30ed\u30a2\u4e9c\u79d1(Anomochlooideae)<br \/>\n\u3000\u3000\u3000\u2514\u252c\u30d5\u30a1\u30eb\u30b9\u4e9c\u79d1(Pharoideae)<br \/>\n\u3000\u3000\u3000\u3000\u2514\u252c\u30d7\u30a8\u30ea\u30a2\u4e9c\u79d1(Puelioideae)<br \/>\n\u3000\u3000\u3000\u3000\u3000\u2514<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30a4\u30cd\u79d1(Poaceae)<\/span> \u2500\u2192 \u3010<a href=\"#Poaceae_Phylogeny2\">\u30a4\u30cd\u79d1<\/a>\u3011<br \/>\n\u3000\u3000\u3000\u3000\u3000\u3000\u2514\u252cBEP\u5206\u5c90\u7fa4(BEP clade)<br \/>\n\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2514PACCMAD\u5206\u5c90\u7fa4(PACCMAD clade)\n<\/div>\n\n\n--><\/p>\n<p><a href=\"#Top\">To Top<\/a><\/p>\n<hr id=\"Poaceae_Phylogeny\">\n\u3010\u30a4\u30cd\u79d1(Poaceae)\u306e\u7cfb\u7d71\u6982\u8981\u3011<a href=\"#Huang_2022\">Huang <i>et al<\/i>.(2022)<\/a><br \/>\n<span style=\"color:green; background-color:inherit; font-size:small;\">\u3000\u30a4\u30cd\u79d1\u306f\u3001\u7d0411000\u7a2e\u304b\u3089\u69cb\u6210\u3055\u308c\u308b\u88ab\u5b50\u690d\u7269\u3068\u3057\u3066\u6700\u5927\u306e\u79d1\u3067\u3059\u3002\u9ad8\u6e29\u3001\u4e7e\u71e5\u3001\u4f4e\u4e8c\u9178\u5316\u70ad\u7d20\u6fc3\u5ea6\u3067\u3082\u751f\u80b2\u3059\u308b\u3053\u3068\u304c\u3067\u304d\u308bC4\u5149\u5408\u6210\u306b\u9069\u5fdc\u3057\u305f\u7a2e\u304c\u542b\u307e\u308c\u3066\u3044\u3066\u3001\u30a4\u30cd(C3)\u3001\u30b3\u30e0\u30ae(C3)\u3001\u30c8\u30a6\u30e2\u30ed\u30b3\u30b7(C4)\u306a\u3069\u91cd\u8981\u7a40\u7269\u3082\u591a\u6570\u542b\u307e\u308c\u3066\u3044\u307e\u3059\u3002\u5149\u5408\u6210\u30bf\u30a4\u30d7\u306b\u304a\u3051\u308bC3\/C4\u306e\u5225\u306f\u3001<a href=\"#GPWG_2012\">GPWG-II(2012)<\/a>\u3001<a href=\"#Takeda_1985\">\u6b66\u7530\u3089(1985)<\/a>, <a href=\"#Yoshimura_2015\">\u5409\u6751(2015)<\/a>\u7b49\u3092\u53c2\u8003\u306b\u3057\u3066\u793a\u3057\u3066\u3044\u307e\u3059\u3002<\/span><\/p>\n<div style=\"line-height: 1em; font-family: monospace; padding:0 0 1em 0;\">\n\u3000\u3000\u3000\u3000\u250cPACMAD\u5206\u5c90\u7fa4(PACMAD Clade)<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u3000\u3000\u3000\u250c<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Cynodon\"><span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30d2\u30b2\u30b7\u30d0\u4e9c\u79d1(Chloridoideae)<\/span><\/a>> \u2500\u2192 \u3010<a href=\"#Chloridoideae_Phylogeny\">\u30d2\u30b2\u30b7\u30d0\u4e9c\u79d1<\/a>\u3011<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u3000\u3000\u3000\u2502\u2502\u3000\u250c\u30aa\u30d2\u30b2\u30b7\u30d0\u9023(Cynodonteae) C4<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u3000\u3000\u250c\u2524\u2502\u250c\u253c\u30b7\u30d0\u9023(Zoysieae) C3<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u3000\u3000\u2502\u2502\u2514\u2524\u251c\u30b9\u30ba\u30e1\u30ac\u30e4\u9023(Eragrostideae) C4<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u3000\u250c\u2524\u2502\u3000\u2502\u2514\u30c8\u30ea\u30e9\u30d5\u30a3\u30b9\u9023(Triraphideae)<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u3000\u2502\u2502\u2502\u3000\u2514\u30bb\u30f3\u30c8\u30ed\u30dd\u30c7\u30a3\u30a2\u9023(Centropodieae) C3\/C4<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u3000\u2502\u2502\u2514<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30c0\u30f3\u30bd\u30cb\u30a2\u4e9c\u79d1(Danthonioideae)<\/span><br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u250c\u2524\u2514<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Arundo\"><span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30c0\u30f3\u30c1\u30af\u4e9c\u79d1(Arundinoideae)<\/span><\/a> C3<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u2502\u2502\u250c<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Chasmanthium\"><span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30ad\u30d3\u4e9c\u79d1(Panicoideae)<\/span><\/a> \u2500\u2192 \u3010<a href=\"#Panicoideae_Phylogeny\">\u30ad\u30d3\u4e9c\u79d1<\/a>\u3011<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u2502\u2502\u2502\u2502\u3000\u3000\u3000\u3000\u3000\u3000\u250c\u30a6\u30b7\u30af\u30b5\u9023(Andropogoneae) C4<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u2502\u2514\u2524\u2502\u3000\u3000\u3000\u3000\u250c\u252c\u2534\u30c8\u30c0\u30b7\u30d0\u9023(Arundinelleae) C4<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u2502\u3000\u2502\u2502\u3000\u3000\u3000\u250c\u2524\u2514\u30b9\u30ba\u30e1\u30ce\u30d2\u30a8\u9023(Paspaleae) C4<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2514\u2524\u3000\u2502\u2502\u3000\u3000\u250c\u2524\u2514\u30ad\u30d3\u9023(Paniceae) C3\/C4<br \/>\n\u3000\u3000\u3000\u3000\u2502\u3000\u2502\u3000\u2502\u2502\u3000\u250c\u2524\u2514\u30c8\u30ea\u30b9\u30bf\u30ad\u30a2\u9023(Tristachyideae) C4<br \/>\n\u3000\u3000\u3000\u3000\u2502\u3000\u2502\u3000\u2502\u2502\u250c\u2524\u2514\u30ae\u30cd\u30ea\u30a6\u30e0\u9023(Gynerieae)<br \/>\n\u3000\u3000\u3000\u250c\u2524\u3000\u2502\u3000\u2502\u2514\u2524\u2514\u252c\u30c1\u30b5\u30ce\u30ec\u30ca\u9023(Thysanoiaeneae)<br \/>\n\u3000\u3000\u3000\u2502\u2502\u3000\u2502\u3000\u2502\u3000\u2502\u3000\u2514\u30bb\u30f3\u30c8\u30c6\u30ab\u9023(Centotheceae) C3<br \/>\n\u3000\u3000\u3000\u2502\u2502\u3000\u2502\u3000\u2502\u3000\u2514\u252c\u30bc\u30ae\u30c6\u30b9\u9023(Zuegiteae)<br \/>\n\u3000\u3000\u250c\u2524\u2502\u3000\u2502\u3000\u2502\u3000\u3000\u2514\u30c1\u30e3\u30b9\u30de\u30f3\u30c6\u30a3\u30a6\u30e0\u9023(Chasmanthieae)<br \/>\n\u3000\u3000\u2502\u2502\u2502\u3000\u2502\u3000\u2514<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30df\u30af\u30e9\u30a4\u30e9\u4e9c\u79d1(Micrairoideae)<\/span><br \/>\n\u3000\u250c\u2524\u2502\u2502\u3000\u2502\u3000\u3000\u2514\u252c\u30a4\u30b5\u30af\u30cd\u9023(Isachneae) C3<br \/>\n\u3000\u2502\u2502\u2502\u2502\u3000\u2502\u3000\u3000\u3000\u2514\u30a8\u30ea\u30a2\u30af\u30cd\u9023(Eriachneae) C4<br \/>\n\u3000\u2502\u2502\u2502\u2502\u3000\u2514<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30a2\u30ea\u30b9\u30c6\u30a3\u30c0\u4e9c\u79d1(Aristidoideae)<\/span><br \/>\n\u250c\u2524\u2502\u2502\u2502\u3000\u3000\u3000\u2514\u30a2\u30ea\u30b9\u30c6\u30a3\u30c0\u9023(Aristideae) C4<br \/>\n\u2502\u2502\u2502\u2502\u2514BEP\u5206\u5c90\u7fa4(BEP Clade) C3<br \/>\n\u2502\u2502\u2502\u2502\u3000\u2502\u250c\u252c<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Poa\"><span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30a4\u30c1\u30b4\u30c4\u30ca\u30ae\u4e9c\u79d1(Pooideae)<\/span><\/a> C3<br \/>\n\u2524\u2502\u2502\u2502\u3000\u2514\u2524\u2514<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Shibataea\"><span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30bf\u30b1\u4e9c\u79d1(Bambusoideae)<\/span><\/a> C3 \u2500\u2192 \u3010<a href=\"#Bambusoideae_Phylogeny\">\u30bf\u30b1\u4e9c\u79d1<\/a>\u3011<br \/>\n\u2502\u2502\u2502\u2502\u3000\u3000\u2514<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Oryza\"><span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30a4\u30cd\u4e9c\u79d1(Orizoideae)<\/span><\/a>=\u30a8\u30fc\u30eb\u30cf\u30eb\u30bf\u4e9c\u79d1(Ehrhartoideae) C3<br \/>\n\u2502\u2502\u2502\u2514\u30d7\u30a8\u30ea\u30a2\u9023(Atractocarpeae)\u2500\u2500\u2500\u252c<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30d7\u30a8\u30ea\u30a2\u4e9c\u79d1<\/span><br \/>\n\u2502\u2502\u2514\u30b0\u30a2\u30c7\u30e5\u30a8\u30e9\u9023(Guaduelleae) \u2500\u2500\u2500\u2518<span style=\"color:maroon; background-color:inherit; font-weight:700;\">(Puelioideae)<\/span><br \/>\n\u2502\u2514<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30d5\u30a1\u30eb\u30b9\u4e9c\u79d1(Pharoideae)<\/span><br \/>\n\u2514<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30a2\u30ce\u30e2\u30af\u30ed\u30a2\u4e9c\u79d1(Anomochlooideae)<\/span>\n<\/div>\n<p><a href=\"#Top\">To Top<\/a><\/p>\n<hr id=\"Poaceae_Phylogeny_GPWG-II\">\n\u3010\u30a4\u30cd\u79d1(Poaceae)\u306e\u7cfb\u7d71\u6982\u8981\u3011<a href=\"#GPWG_2012\">GPWG-II(2012)<\/a><br \/>\n<span style=\"color:green; background-color:inherit; font-size:small;\">\u3000\u5149\u5408\u6210\u30bf\u30a4\u30d7\u306b\u304a\u3051\u308bC3\/C4\u306e\u5225\u306f\u3001<a href=\"#GPWG_2012\">GPWG-II(2012)<\/a>\u3001<a href=\"#Takeda_1985\">\u6b66\u7530\u3089(1985)<\/a>, <a href=\"#Yoshimura_2015\">\u5409\u6751(2015)<\/a>\u7b49\u3092\u53c2\u8003\u306b\u3057\u3066\u793a\u3057\u3066\u3044\u307e\u3059\u3002<\/span><\/p>\n<div style=\"line-height: 1em; font-family: monospace; padding:0 0 1em 0;\">\n\u3000\u3000\u3000\u3000\u3000\u250c\u30ad\u30d3\u4e9c\u79d1(Panicoideae)\uff1a250\u5c5e32366\u7a2e C3\/C4 \u2500\u2192 \u3010<a href=\"#Panicoideae_Phylogeny\">\u30ad\u30d3\u4e9c\u79d1<\/a>\u3011<br \/>\n\u3000\u3000\u3000\u3000\u250c\u2524\u3000\u250c\u30df\u30af\u30e9\u30a4\u30e9\u4e9c\u79d1(Micrairoideae)\uff1a12\u5c5e188\u7a2e C3\/C4<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u250c\u2534\u30c0\u30f3\u30c1\u30af\u4e9c\u79d1(Arundinoideae)\uff1a9\u5c5e45\u7a2e C3<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2514\u2524\u250c\u30c0\u30f3\u30bd\u30cb\u30a2\u4e9c\u79d1(Danthonioideae)\uff1a26\u5c5e281\u7a2e<br \/>\n\u3000\u3000\u3000\u250c\u2524\u3000\u2514\u2534\u252c\u30bb\u30f3\u30c8\u30ed\u30dd\u30c7\u30a3\u30a2\u5206\u5c90\u7fa4(Centropodie clade)\uff1a3\u5c5e6\u7a2e C3\/C4<br \/>\n\u3000\u3000\u250c\u2524\u2502\u3000\u3000\u3000\u2514\u30d2\u30b2\u30b7\u30d0\u4e9c\u79d1(Chloridoideae)\uff1a125\u5c5e1601\u7a2e C3\/C4 \u2500\u2192 \u3010<a href=\"#Chloridoideae_Phylogeny\">\u30d2\u30b2\u30b7\u30d0\u4e9c\u79d1<\/a>\u3011<br \/>\n\u3000\u3000\u2502\u2502\u2514\u30a2\u30ea\u30b9\u30c6\u30a3\u30c0\u4e9c\u79d1(Aristidoideae)\uff1a27\u5c5e349\u7a2e C4<br \/>\n\u3000\u250c\u2524\u2514\u252c\u30a8\u30fc\u30eb\u30cf\u30eb\u30bf\u4e9c\u79d1(Ehrhartoideae)\uff1a11\u5c5e111\u7a2e = \u30a4\u30cd\u4e9c\u79d1(Orizoideae) C3 \u2500\u2192 \u3010<a href=\"#Poaceae_Phylogeny\">\u30a4\u30cd\u4e9c\u79d1<\/a>\u3011<br \/>\n\u250c\u2524\u2502\u3000\u2514\u252c\u30bf\u30b1\u4e9c\u79d1(Bambusoideae)\uff1a16\u5c5e1462\u7a2e C3\u2500\u2192 \u3010<a href=\"#Bambusoideae_Phylogeny\">\u30bf\u30b1\u4e9c\u79d1<\/a>\u3011<br \/>\n\u2524\u2502\u2502\u3000\u3000\u2514\u30a4\u30c1\u30b4\u30c4\u30ca\u30ae\u4e9c\u79d1(Pooideae)\uff1a48\u5c5e3850\u7a2e C3<br \/>\n\u2502\u2502\u2514\u30d7\u30a8\u30ea\u30a2\u4e9c\u79d1(Puelioideae)\uff1a1\u5c5e11\u7a2e<br \/>\n\u2502\u2514\u30d5\u30a1\u30eb\u30b9\u4e9c\u79d1(Pharoideae)\uff1a1\u5c5e13\u7a2e<br \/>\n\u2514\u30a2\u30ce\u30e2\u30af\u30ed\u30a2\u4e9c\u79d1(Anomochlooideae)\uff1a2\u5c5e4\u7a2e\n<\/div>\n<p><a href=\"#Top\">To Top<\/a><\/p>\n<hr id=\"Chloridoideae_Phylogeny\">\n\u3010\u30d2\u30b2\u30b7\u30d0\u4e9c\u79d1(Chloridoideae)\u5468\u8fba\u306e\u7cfb\u7d71\u6982\u8981\u3011<a href=\"#Wang_2022\">Wang <i>et al<\/i>.(2022)<\/a><br \/>\n<br \/>\n<span style=\"color:green; background-color:inherit; font-size:small;\">\u3000\u30d2\u30b2\u30b7\u30d0\u4e9c\u79d1\u306f140\u5c5e1400\u7a2e\u4ee5\u4e0a\u3067\u69cb\u6210\u3055\u308c(<a href=\"#Wang_2022\">Wang <i>et al.<\/i>.2022<\/a>)\u3001\u305d\u306e\u3046\u3061\u7d04\u534a\u6570\u306e\u7a2e\u304c\u30aa\u30d2\u30b2\u30b7\u30d0\u9023\u306e\u6240\u5c5e\u3067\u3059\u3002C4\u5149\u5408\u6210\u306b\u5bfe\u5fdc\u3057\u305f\u7a2e\u304c\u591a\u304f\u542b\u307e\u308c\u3066\u3044\u308b\u3053\u3068\u304b\u3089\u3001\u8fd1\u672a\u6765\u306e\u98df\u7ce7\u990c\u6599\u3068\u3057\u3066\u3082\u671f\u5f85\u3055\u308c\u3066\u3044\u307e\u3059\u3002\u5149\u5408\u6210\u30bf\u30a4\u30d7\u306b\u304a\u3051\u308bC3\/C4\u306e\u5225\u306f\u3001<a href=\"#GPWG_2012\">GPWG-II(2012)<\/a>\u3001<a href=\"#Takeda_1985\">\u6b66\u7530\u3089(1985)<\/a>, <a href=\"#Yoshimura_2015\">\u5409\u6751(2015)<\/a>\u7b49\u3092\u53c2\u8003\u306b\u3057\u3066\u793a\u3057\u3066\u3044\u307e\u3059\u3002<\/span><\/p>\n<div style=\"line-height: 1em; font-family: monospace; padding:0 0 1em 0;\">\n\u250c<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30d2\u30b2\u30b7\u30d0\u4e9c\u79d1(Chloridoideae)<\/span><br \/>\n\u2502\u2502\u3000\u3000\u3000\u250c\u30aa\u30d2\u30b2\u30b7\u30d0\u9023(Cynodonteae) C4<br \/>\n\u2524\u2502\u3000\u3000\u3000\u2502\u2502\u3000\u3000\u3000\u3000\u3000\u250c\u30c8\u30ea\u30aa\u30c7\u30a3\u30a2\u4e9c\u9023(Triodiinae)<br \/>\n\u2502\u2502\u3000\u3000\u3000\u2502\u2502\u3000\u3000\u3000\u250c\u252c\u2534\u30aa\u30ea\u30ca\u30b9\u5c5e(Orinus)<br \/>\n\u2502\u2502\u3000\u3000\u3000\u2502\u2502\u3000\u3000\u250c\u2524\u2514\u30af\u30ec\u30a4\u30b9\u30c8\u30b8\u30a7\u30ca\u30b9\u5c5e(Cleistogenes)<br \/>\n\u2502\u2502\u3000\u3000\u3000\u2502\u2502\u3000\u250c\u2524\u2514\u252c\u30dc\u30a6\u30c6\u30ed\u30a6\u30a2\u4e0a\u4e9c\u9023(supersubtribe Gouiniodinae)<br \/>\n\u2502\u2502\u3000\u3000\u3000\u2502\u2502\u3000\u2502\u2502\u3000\u2514\u30bf\u30c4\u30ce\u30c4\u30e1\u30ac\u30e4\u4e9c\u9023(Dactylocteniinae)<br \/>\n\u2502\u2502\u3000\u3000\u3000\u2502\u2502\u250c\u2524\u2514\u252c\u30a8\u30ec\u30a6\u30b7\u30f3\u4e9c\u9023(Eleusininae)<br \/>\n\u2502\u2502\u3000\u3000\u3000\u2502\u2514\u2524\u2502\u3000\u2514\u30a2\u30a8\u30eb\u30ed\u30d7\u30b9\u4e9c\u9023(Aeluropodinae)<br \/>\n\u2502\u2502\u3000\u3000\u250c\u2524\u3000\u2502\u2514\u30c8\u30ea\u30dd\u30b4\u30f3\u4e9c\u9023(tripogoninae)<br \/>\n\u2502\u2502\u3000\u250c\u2524\u2502\u3000\u2514\u30dc\u30a6\u30c6\u30ed\u30a6\u30a2\u4e0a\u4e9c\u9023(supersubtribe Boutelouodinae)<br \/>\n\u2502\u2502\u250c\u2524\u2502\u2514\u30b7\u30d0\u9023(Zoysieae) C3<br \/>\n\u2502\u2514\u2524\u2502\u2514\u30b9\u30ba\u30e1\u30ac\u30e4\u9023(Eragrostideae) C4<br \/>\n\u2502\u3000\u2502\u2514\u30c8\u30ea\u30e9\u30d5\u30a3\u30b9\u9023(Triraphideae)<br \/>\n\u2502\u3000\u2514\u30bb\u30f3\u30c8\u30ed\u30dd\u30c7\u30a3\u30a2\u9023(Centropodieae) C3\/C4<br \/>\n\u2514\u5916\u7fa4(outgroup)\uff1a\u30c0\u30f3\u30bd\u30cb\u30a2\u4e9c\u79d1<br \/>\n\u3000\u2502\u3000\u250c\u30ab\u30ea\u30d5\u30a9\u30eb\u30cb\u30a2\u30aa\u30fc\u30c8\u30b0\u30e9\u30b9 <i>Danthonia callifornica<\/i><br \/>\n\u3000\u2514\u252c\u2534\u30b7\u30ed\u30ac\u30cd\u30e8\u30b7<i>Cortaderia selloana<\/i><br \/>\n\u3000\u3000\u2514\u30ad\u30aa\u30ce\u30af\u30ed\u30a2\u30fb\u30de\u30af\u30e9<i>Chionochloa macra<\/i>\n<\/div>\n<p><a href=\"#Top\">To Top<\/a><\/p>\n<hr id=\"Panicoideae_Phylogeny\">\n\u3010\u30ad\u30d3\u4e9c\u79d1(Panicoideae)\u5468\u8fba\u306e\u7cfb\u7d71\u6982\u8981\u3011<a href=\"#Burke_2016\">Burke <i>et al<\/i>.(2016)<\/a><br \/>\n<br \/>\n<span style=\"color:green; background-color:inherit; font-size:small;\">\u3000\u30c8\u30a6\u30e2\u30ed\u30b3\u30b7\u3001\u7a17\u3001\u7c9f\u306a\u3069\u91cd\u8981\u306a\u7a40\u7269\u304c\u591a\u6570\u542b\u307e\u308c\u3066\u3044\u308b\u30ad\u30d3\u4e9c\u79d1\u3067\u306f\u3001212\u5c5e3316\u7a2e\u304c\u8a18\u8f09\u3055\u308c\u3066\u3044\u308b\u3068\u3055\u308c\u3001\u4e9c\u79d1\u5185\u306b\u306f\u5c11\u306a\u304f\u3068\u308210\u306e\u9023\u304c\u8b58\u5225\u3055\u308c\u3066\u3044\u308b\u69d8\u3067\u3059\u3002\u5149\u5408\u6210\u30bf\u30a4\u30d7\u306b\u304a\u3051\u308bC3\/C4\u306e\u5225\u306f\u3001<a href=\"#GPWG_2012\">GPWG-II(2012)<\/a>\u3001<a href=\"#Takeda_1985\">\u6b66\u7530\u3089(1985)<\/a>, <a href=\"#Yoshimura_2015\">\u5409\u6751(2015)<\/a>\u7b49\u3092\u53c2\u8003\u306b\u3057\u3066\u793a\u3057\u3066\u3044\u307e\u3059\u3002<\/span><\/p>\n<div style=\"line-height: 1em; font-family: monospace; padding:0 0 1em 0;\">\n\u3000\u3000\u3000\u3000\u250c<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30ad\u30d3\u4e9c\u79d1(Panicoides)<\/span><br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u3000\u3000\u3000\u3000\u250c\u30a6\u30b7\u30af\u30b5\u9023(Andropogoneae) C4<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u3000\u3000\u250c\u252c\u2534\u30c8\u30c0\u30b7\u30d0\u9023(Arundinelleae) C4<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u3000\u250c\u2524\u2514\u30b9\u30ba\u30e1\u30ce\u30d2\u30a8\u9023(Paspaleae) C4<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2502\u250c\u2524\u2514\u30ad\u30d3\u9023(Paniceae) C3\/C4<br \/>\n\u3000\u3000\u3000\u3000\u2502\u2514\u2524\u2514\u30b9\u30ba\u30e1\u30ce\u30d2\u30a8\u9023(Paspaleae) C4<br \/>\n\u3000\u3000\u3000\u250c\u2524\u3000\u2502\u3000\u250c\u252c\u30bb\u30f3\u30c8\u30c6\u30ab\u9023(Centotheceae) C3<br \/>\n\u3000\u3000\u3000\u2502\u2502\u3000\u2502\u250c\u2524\u2514\u30c1\u30b5\u30ce\u30ec\u30ca\u9023(Thysanoiaeneae)<br \/>\n\u3000\u3000\u3000\u2502\u2502\u3000\u2514\u2524\u2514\u30c8\u30ea\u30b9\u30bf\u30ad\u30a2\u9023(Tristachyideae) C4<br \/>\n\u3000\u3000\u3000\u2502\u2502\u3000\u3000\u2514\u252c\u30c1\u30e3\u30b9\u30de\u30f3\u30c6\u30a3\u30a6\u30e0\u9023(Chasmanthieae)<br \/>\n\u3000\u3000\u250c\u2524\u2502\u3000\u3000\u3000\u2514\u30bc\u30ae\u30c6\u30b9\u9023(Zuegiteae)<br \/>\n\u3000\u3000\u2502\u2502\u2502\u3000\u3000\u250c\u30d2\u30b2\u30b7\u30d0\u4e9c\u79d1(Chloridoideae)<br \/>\n\u3000\u3000\u2502\u2502\u2502\u250c\u252c\u2534\u30c0\u30f3\u30c8\u30cb\u30a2\u4e9c\u79d1(Danthonoideae)<br \/>\n\u3000\u250c\u2524\u2502\u2514\u2524\u2514\u30df\u30af\u30e9\u30a4\u30e9\u79d1(Micrairoideae) C3\/C4<br \/>\n\u3000\u2502\u2502\u2502\u3000\u2514\u30a2\u30ea\u30b9\u30c6\u30a3\u30c0\u4e9c\u79d1(Aristidoideae)<br \/>\n\u250c\u2524\u2502\u2502\u3000\u250c\u30a4\u30c1\u30b4\u30c4\u30ca\u30ae\u4e9c\u79d1(Pooideae) C3<br \/>\n\u2502\u2502\u2502\u2514\u252c\u2534\u30bf\u30b1\u4e9c\u79d1(Bambusoideae) C3<br \/>\n\u2524\u2502\u2502\u3000\u2514\u30a4\u30cd\u4e9c\u79d1(Orizoideae) C3<br \/>\n\u2502\u2502\u2514\u30d7\u30a8\u30ea\u30a2\u4e9c\u79d1(Puelioideae)<br \/>\n\u2502\u2514\u30d5\u30a1\u30eb\u30b9\u4e9c\u79d1(Pharoideae)<br \/>\n\u2514\u30a2\u30ce\u30e2\u30af\u30ed\u30a2\u4e9c\u79d1(Anomochlooideae)\n<\/div>\n<p><a href=\"#Top\">To Top<\/a><\/p>\n<hr id=\"Bambusoideae_Phylogeny\">\n\u3010\u30bf\u30b1\u4e9c\u79d1(Bambusoideae)\u306e\u7cfb\u7d71\u6982\u8981\u3011<a href=\"#Wysocki_2015\">Wysocki <i>et al<\/i>.(2015)<\/a><br \/>\n<span style=\"color:green; background-color:inherit; font-size:small;\">\u3000\u30bf\u30b1\u4e9c\u79d1\u306f\u3001115\u5c5e1450\u7a2e\u304c\u8a18\u8f09\u3055\u308c\u3066\u3044\u3066\u3001\u5c11\u306a\u304f\u3068\u30823\u3064\u306e\u9023\u306b\u76f8\u5f53\u3059\u308b\u30af\u30ec\u30fc\u30c9(\u5206\u5c90\u7fa4)\u304c\u8b58\u5225\u3067\u304d\u308b\u3088\u3046\u3067\u3059\u3002\u3001<\/span><\/p>\n<div style=\"line-height: 1em; font-family: monospace; padding:0 0 1em 0;\">\n\u3000\u250c\u30bf\u30b1\u4e9c\u79d1(Bambusoideae)<br \/>\n\u3000\u2502\u2502\u3000\u250c<a href=\"#Bambusoideae_Phylogeny\">\u30bf\u30b1\u9023(Bambiseae)<\/a> \u2500\u2192 \u3010<a href=\"#Bambuseae_Phylogeny\">\u30bf\u30b1\u9023<\/a>\u3011<br \/>\n\u250c\u2524\u2502\u250c\u2524\u2514\u252c\u65e7\u71b1\u5e2f\u533a\u306e\u30bf\u30b1\u9023(Paleotropical)<br \/>\n\u2502\u2502\u2514\u2524\u2502\u3000\u2514\u65b0\u71b1\u5e2f\u533a\u306e\u30bf\u30b1\u9023(Neotropical)<br \/>\n\u2524\u2502\u3000\u2502\u2514\u9ecd\u7afa(\u3057\u3087\u3058\u304f)\u9023(Olyreae)<br \/>\n\u2502\u2502\u3000\u2514<a href=\"#Arundinarieae_Phylogeny\">\u30e1\u30c0\u30b1\u9023(Arundinarieae)<\/a> \u2500\u2192 \u3010<a href=\"#Arundinarieae_Phylogeny\">\u30e1\u30c0\u30b1\u9023<\/a>\u3011<br \/>\n\u2502\u2514<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Lolium\">\u30db\u30bd\u30e0\u30ae <i>Lolium perenne<\/i><\/a>\uff1a\u30a4\u30c1\u30b4\u30c4\u30ca\u30ae\u4e9c\u79d1(Pooideae)<br \/>\n\u2514\u30a2\u30e1\u30ea\u30ab\u30de\u30b3\u30e2 <i>Zizania aquatica<\/i>\uff1a\u30a8\u30fc\u30eb\u30cf\u30eb\u30bf\u4e9c\u79d1(Ehrhartoideae)<br \/>\n\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000=\u30a4\u30cd\u4e9c\u79d1(Orizoideae)\n<\/div>\n<p><a href=\"#Top\">To Top<\/a><\/p>\n<hr id=\"Bambuseae_Phylogeny\">\n\u3010\u30bf\u30b1\u9023(Bambuseae)\u5468\u8fba\u306e\u7cfb\u7d71\u6982\u8981\u3011<a href=\"#Triplett_2010\">Triplett <i>et al<\/i>.(2010)<\/a><br \/>\n<span style=\"color:green; background-color:inherit; font-size:small;\"><br \/>\n\u3000\u30b9\u30c8\u30ec\u30d7\u30c8\u30b8\u30fc\u30ca\u9023\u306f\u3001\u30bf\u30b1\u4e9c\u79d1\u3068\u306e\u95a2\u4fc2\u304c\u6307\u6458\u3055\u308c\u305f\u3053\u3068\u3082\u3042\u308a\u307e\u3057\u305f\u304c<a href=\"#Iioka_1958\">(\u98ef\u5ca1,1958)<\/a>\u3001\u73fe\u5728\u3067\u306f\u30a4\u30cd\u4e9c\u79d1\u5185\u306e4\u3064\u76ee\u306e\u9023\u3068\u3055\u308c\u3066\u3044\u307e\u3059(<a href=\"#Saarela_2018\">Saarela <i>et al.<\/i>,2018)<\/a><\/span><\/p>\n<div style=\"line-height: 1em; font-family: monospace; padding:0 0 1em 0;\">\n\u250c\u30d5\u30a1\u30eb\u30b9\u4e9c\u79d1(Pharoideae)<br \/>\n\u2524\u250c\u252c\u30a8\u30fc\u30eb\u30cf\u30eb\u30bf\u4e9c\u79d1(\u30a4\u30cd\u4e9c\u79d1)(Ehrhartoideae)<br \/>\n\u2502\u2502\u2514\u30b9\u30c8\u30ec\u30d7\u30c8\u30b8\u30fc\u30ca\u9023(Streptogyneae)<br \/>\n\u2514\u2524\u250c\u30a4\u30c1\u30b4\u30c4\u30ca\u30ae\u4e9c\u79d1(Pooideae)<br \/>\n\u3000\u2514\u2534<span style=\"color:maroon; background-color:inherit; font-weight:700;\">\u30bf\u30b1\u4e9c\u79d1(Bambusoideae)<\/span><br \/>\n\u3000\u3000\u3000\u2502\u3000\u250c(\u71b1\u5e2f\u6728\u6027\u30d0\u30f3\u30d6\u30fc:Tropical Woody Bamboos)<br \/>\n\u3000\u3000\u3000\u2502\u250c\u2534(\u8349\u6027\u30d0\u30f3\u30d6\u30fc:Herbaceous Bamboos)<br \/>\n\u3000\u3000\u3000\u2514\u2534(\u6e29\u5e2f\u6728\u6027\u30d0\u30f3\u30d6\u30fc:Temperate Woody Bamboos)<br \/>\n\u3000\u3000\u3000\u3000\u3000\u2502\u250c\u30bf\u30e0\u30ce\u30ab\u30e9\u30e0\u30b9\u985e(I. Bergbamboes)<br \/>\n\u3000\u3000\u3000\u3000\u3000\u2514\u2524\u250c\u30a2\u30d5\u30ea\u30ab\u9ad8\u5c71\u6027\u30d0\u30f3\u30d6\u30fc(II. African apline bamboos)<br \/>\n\u3000\u3000\u3000\u3000\u3000\u3000\u2514\u253c\u9999\u7af9\u5206\u5c90\u7fa4(III. Chimonocalamus)<br \/>\n\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u251c<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Shibataea\">\u30aa\u30ab\u30e1\u30b6\u30b5\u5206\u5c90\u7fa4(IV. Shibataeae clade)<\/a><br \/>\n\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u251c<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Phyllostachys\">\u30de\u30c0\u30b1\u5206\u5c90\u7fa4(V. Phyllostachys clade)<\/a><br \/>\n\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2514<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Sasaella\">\u30a2\u30ba\u30de\u30b6\u30b5\u5206\u5c90\u7fa4(VI. Arundinaria clade)<\/a><br \/>\n\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2514\u252c(Sasa-Arundinaria <i>sensu Stricto<\/i> subclade)<br \/>\n\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2514\u252c(Sinicae subclade)<br \/>\n\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2514(Medake subclade)\n<\/div>\n<p><a href=\"#Top\">To Top<\/a><\/p>\n<hr id=\"Arundinarieae_Phylogeny\">\n\u3010\u30e1\u30c0\u30b1\u9023(Arundinarieae)\u5468\u8fba\u306e\u7cfb\u7d71\u6982\u8981\u3011<a href=\"#Attigala_2016\">Attigala <i>et al<\/i>.(2016)<\/a><br \/>\n<span style=\"color:green; background-color:inherit; font-size:small;\">\u30e1\u30c0\u30b1\u9023\u5185\u306b12\u306e\u30af\u30ec\u30fc\u30c9\u304c\u8b58\u5225\u3055\u308c\u3066\u3044\u307e\u3059\u3002<\/span><\/p>\n<div style=\"line-height: 1em; font-family: monospace; padding:0 0 1em 0;\">\n\u250c\u30a4\u30c1\u30b4\u30c4\u30ca\u30ae\u4e9c\u79d1(Pooideae)\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000 \u2510<br \/>\n\u2524\u2514\u252c<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Lolium\">\u30db\u30bd\u30e0\u30ae <i>Lolium perenne<\/i><\/a>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2502<br \/>\n\u2502\u3000\u2514<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Hordeum\">\u56db\u6761\u30aa\u30aa\u30e0\u30ae <i>Hordeum vulgare<\/i> Subsp. <i>vulgare<\/i><\/a>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2502<br \/>\n\u2502\u250c\u30bf\u30b1\u9023(Bambuseae)\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2502<br \/>\n\u2502\u2502\u2502\u3000\u3000\u250c\u252c\u30a4\u30f3\u30c9\u30c8\u30b2\u30bf\u30b1 <i>Bambusa bambos<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u251c\u5916\u7fa4<br \/>\n\u2514\u2524\u2502\u3000\u250c\u2524\u2514\u30e1\u30ed\u30ab\u30f3\u30ca\u30fb\u30d0\u30af\u30b7\u30d5\u30a7\u30e9 <i>Melocanna baccifera<\/i>\u3000\u3000\u3000\u3000\u3000\u3000 \u3000\u2502(Outgroups)<br \/>\n\u3000\u2502\u2502\u250c\u2524\u2514\u30aa\u30eb\u30e1\u30ab\u30fb\u30ea\u30d5\u30ec\u30af\u30b5 <i>Olmeca reflexa<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2502<br \/>\n\u3000\u2502\u2514\u2524\u2514\u30c1\u30e5\u30b9\u30b1\u30a2\u30fb\u30b9\u30da\u30af\u30bf\u30d3\u30ea\u30b9 <i>Chusquea spectabilis<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2502<br \/>\n\u3000\u2502\u3000\u2514\u252c\u30aa\u30aa\u30d0\u30c1\u30c2\u30df\u30b6\u30b5 <i>Olyra latifolia<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000 \u2502<br \/>\n\u3000\u2502\u3000\u3000\u2514\u30af\u30ea\u30d7\u30c8\u30af\u30ed\u30a2\u30fb\u30b9\u30c8\u30ea\u30af\u30c6\u30a3\u30d5\u30a9\u30ea\u30a2 <i>Cryptochloa strictifolia<\/i>\u3000\u2518<br \/>\n\u3000\u2514\u30e1\u30c0\u30b1\u9023(Arundinarieae)<br \/>\n\u3000\u3000\u2502\u3000\u3000\u3000\u3000\u250c\u30a2\u30b7\u30c9\u30b5\u30b5\u30fb\u30d7\u30eb\u30d7\u30ec\u30a2 <i>Acidosasa purpurea<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000 \u2510<br \/>\n\u3000\u3000\u2502\u3000\u3000\u3000\u250c\u2534\u30ab\u30f3\u30b6\u30f3\u30c1\u30af <i>Pleioblastus hindsii<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000 \u2502<br \/>\n\u3000\u3000\u2502\u3000\u3000\u250c\u2524\u3000\u250c\u252c\u30a2\u30eb\u30f3\u30c7\u30a3\u30ca\u30ea\u30a2\u30fb\u30a2\u30d0\u30e9\u30c1\u30a2\u30ca <i>Arundinaria appalachiana<\/i> \u251cVI<br \/>\n\u3000\u3000\u2502\u3000\u3000\u2502\u2502\u250c\u2524\u2514\u30a2\u30eb\u30f3\u30c7\u30a3\u30ca\u30ea\u30a2\u30fb\u30c6\u30af\u30bf <i>Arundinaria tecta<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2502<br \/>\n\u3000\u3000\u2502\u3000\u250c\u2524\u2514\u2524\u2514\u30a2\u30eb\u30f3\u30c7\u30a3\u30ca\u30ea\u30a2\u30fb\u30ae\u30ac\u30f3\u30c6\u30a2 <i>Arundinaria gigentea<\/i>\u3000\u3000\u3000\u3000 \u2502<br \/>\n\u3000\u3000\u2502\u3000\u2502\u2502\u3000\u2514\u30aa\u30aa\u30b5\u30b5 <i>Sasa veitchii<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2518<br \/>\n\u3000\u3000\u2502\u250c\u2524\u2502\u250c\u30a4\u30f3\u30c8\u30ab\u30e9\u30e0\u30b9\u30fb\u30a6\u30a3\u30eb\u30bd\u30cb <i>Indocalamus wilsonii<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000 \u2500VIII<br \/>\n\u3000\u3000\u2502\u2502\u2502\u2514\u2524\u250c\u30d5\u30a7\u30ed\u30ab\u30e9\u30e0\u30b9\u30fb <i>Ferrocalamus rimosivaginus<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000 \u252cIV<br \/>\n\u3000\u3000\u2502\u2502\u2502\u3000\u2514\u2534<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Shibataea\">\u30aa\u30ab\u30e1\u30b6\u30b5 <i>Shibataea kumsaca<\/i><\/a>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2518<br \/>\n\u3000\u3000\u2514\u2524\u2502\u3000\u3000\u250c\u30d5\u30a1\u30fc\u30b2\u30b7\u30a2\u30fb\u30cb\u30c1\u30c0 <i>Fargesia nitida<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u252cV<br \/>\n\u3000\u3000\u3000\u2502\u2502\u3000\u250c\u2534<a href=\"https:\/\/ichihashi.me\/wp\/?p=40640#Phyllostachys\">\u30db\u30c6\u30a4\u30c1\u30af <i>Phyllostachys aurea<\/i><\/a>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000 \u3000 \u2518<br \/>\n\u3000\u3000\u3000\u2502\u2502\u3000\u251c\u6c34\u94f6\u7af9 <i>Indocalamus sinicus<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2500X<br \/>\n\u3000\u3000\u3000\u2502\u2502\u250c\u2524\u3000\u250c\u9999\u7af9\u5c5e\u306e\u4e00\u7a2e <i>Chimonocal\uff41mus<\/i> sp.\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2500III<br \/>\n\u3000\u3000\u3000\u2502\u2502\u2502\u2502\u250c\u2534\u252c\u30af\u30eb\u30ca\u30fb\u30c7\u30f3\u30b7\u30d5\u30a9\u30ea\u30a2 <i>Kuruna densifolia<\/i>\u3000\u3000\u3000\u3000\u3000\u3000 \u3000 \u252cXII<br \/>\n\u3000\u3000\u3000\u2502\u2514\u2524\u2514\u2524\u3000\u2514\u30af\u30eb\u30ca\u30fb\u30c7\u30d3\u30ea\u30b9 <i>Kuruna debilis<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000 \u2518<br \/>\n\u3000\u3000\u3000\u2502\u3000\u2502\u3000\u251c\u30aa\u30eb\u30c7\u30a2\u30cb\u30a2\u30fb\u30a2\u30eb\u30d4\u30ca <i>Oldeania alpina<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2500II<br \/>\n\u3000\u3000\u3000\u2502\u3000\u2502\u3000\u2514\u9ad8\u9ece\u8d21\u5c71\u7af9(\u304c\u304a\u308a\u3054\u3093\u3061\u304f) <\/rp><\/ruby> <i>Gaoligongshania megalothyrsa<\/i>\u3000\u3000 \u2500IX<br \/>\n\u3000\u3000\u3000\u2502\u3000\u2514\u252c\u7389\u5c71\u7bad\u7af9(\u304e\u3087\u304f\u3056\u3093\u305b\u3093\u3061\u304f) <i>Bergbambos tessellata<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u2500I<br \/>\n\u3000\u3000\u3000\u2502\u3000\u3000\u2514\u30bf\u30e0\u30ce\u30ab\u30e9\u30e0\u30b9\u30fb\u30b9\u30d1\u30c6\u30a3\u30d5\u30ed\u30eb\u30b9 <i>Tamnocalamus sapthiflorus<\/i>\u3000\u3000\u2500VII<br \/>\n\u3000\u3000\u3000\u2514\u30a2\u30c4\u30d0\u30c1\u30af <i>Ampelocalamus calcareus<\/i>\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u3000\u2500XI\n<\/div>\n<p><a href=\"#Top\">To Top<\/a><\/p>\n<hr id=\"Reference\">\n\u3010\u6587\u732e\u3011<br \/>\n<span id=\"GPWG_2001\">GPWG (2001)<\/span> Phylogeny and Subfamilial Classification of the Grasses (Poaceae). Ann Missouri Bot Garden, 88(3), 373-457, <a href=\"hrrps:\/\/doi.org\/10.2307\/3298585\">10.2307\/3298585<\/a>, Accessed: 2025-07-27.<br \/>\n<span id=\"GPGW_2012\">Grass Phylogeny Working Group II<\/a> (2012) New grass phylogeny resolves deep evolutionary relationships and discovers C4 origins, New Phytologist, 193, 304\u2013312, DOI: <a href=\"https:\/doi.org\/10.1111\/j.1469-8137.2011.03972.x\">10.1111\/j.1469-8137.2011.03972.x<\/a>, Accessed: 2025-08-11.<br \/>\n<span id=\"APG-IV\">The Angiosperm Phylogeny Group and others (2016)<\/i> An update of the Angiosperm Phylogeny Group classification for the orders and families of flowering plants: APG IV, Botanical Journal of the Linnean Society, 181(1), 1\u201320, DOI: <a href=\"https:\/\/doi.org\/10.1111\/boj.12385\">10.1111\/boj.12385<\/a>, Accessed: 2023-07-11.<br \/>\n<span id=\"Wang_2022\">Wang R, Zhang X-J, Guo X-X, Xing Y, Qu X-J and Fan S-J (2022)<\/span>, Plastid phylogenomics and morphological character evolution of Chloridoideae (Poaceae), Front Plant Sci, 13:1002724, DOI: <a href=\"https:\/\/doi.org\/10.3389\/fpls.2022.1002724\">10.3389\/fpls.2022.1002724<\/a>, Accessed: 2025-08-11.<br \/>\n<span id=\"Wu_2022\">Wu H, Yang J-B, Liu J-X, Li D-Z and Peng-Fei Ma (2022)<\/span>, Organelle Phylogenomics and Extensive Conflicting Phylogenetic Signals in the Monocot Order Poales, Front Plant Sci, 12, DOI: <a href=\"https:\/\/doi.org\/10.3389\/fpls.2021.824672\">10.3389\/fpls.2021.824672<\/a>, Accessed: 2025-08-11.<br \/>\n<span id=\"Kellogg_2009\">Kellogg EA (2009) The Evolutionary History of Ehrhartoideae, Oryzeae, and <i>Oryza<\/i>, Rice, 2, 1\u201314, DOI <a hre=\"doi.org\/10.1007\/s12284-009-9022-2\">10.1007\/s12284-009-9022-2<\/a>, 2025-08-11.<br \/>\n<span id=\"Huang_2022\">Huang W, Zhang L, Columbus JT, Hu Y, Zhao Y, Tang L, Guo Z, Chen W, McKain M, Bartlett M, Huang C-H, Li D-Z, Ge S and Ma H<\/span> (2022) A well-supported nuclear phylogeny of Poaceae and implications for the evolution of C4 photosynthesis, Molecular Plant, 15, 755\u2013777, DOI: <a href=\"https:\/\/doi.org\/10.1016\/j.molp.2022.01.015\">10.1016\/j.molp.2022.01.015<\/a>, Accessed: 2025-07-27.<br \/>\n<span id=\"Saarela_2018\">Saarela JM, Burke SV, Wysocki WP, Barrett MD, Clark LG, Craine JM, Peterson PM, Soreng RJ, Vorontsova MS and Duvall MR (2018)<\/a>, A 250 plastome phylogeny of the grass family (Poaceae): topological support under different data partitions, . PeerJ, 6:e4299, DOI: <a href=\"https:\/\/doi.org\/10.7717\/peerj.4299\">10.7717\/peerj.4299<\/a>, Accessed: 2025-08-10.<br \/>\n<span id=\"Iioka_1958\">\u98ef\u5ca1\u4e9c\u7dd2 (1958)<\/span> <i>Streptogena<\/i> (\u30a4\u30cd\u79d1)\u306b\u3064\u3044\u3066, \u690d\u7269\u7814\u7a76\u96d1\u8a8c, 33(12), 364-366, Accessed: 2025-07-28.<br \/>\n<span id=\"Burke_2016\">Burke SV, Wysocki WP, Zuloaga FO, Craine JM, Pires JC, Edger PP, Mayfield-Jones D, Clark LG, Kelchner SA and Duvall MR (2016)<\/span> Evolutionary relationships in Panicoid grasses based on plastome phylogenomics (Panicoideae; Poaceae), BMC Plant Biol, 16:140, DOI <a href=\"https:\/\/doi.org\/10.1186\/s12870-016-0823-3\">10.1186\/s12870-016-0823-3<\/a>, Accessed: 2025-07-28.<br \/>\n<span id=\"Wysocki_2015\">Wysocki WP, Clark LG, Attigala L, Ruiz-Sanchez E and Duvall MR (2015<\/span> Evolution of the bamboos (Bambusoideae; Poaceae): a full plastome phylogenomic analysis, BMC Evol Biol, 15:50, DOI: <a href=\"https::\/doi.org\/10.1186\/s12862-015-0321-5\">10.1186\/s12862-015-0321-5<\/a>, accessed: 2025-07-29.<br \/>\n<span id=\"Triplett_2010\">Triplett JK and Clark LG (2010)<\/a> Phylogeny of the Temperate Bamboos (Poaceae: Bambusoideae: Bambuseae) with an Emphasis on <i>Arundinaria<\/i> and Allies, Syst Bot, 35(1). 102\u2013120, DOI: <a href=\"https:\/\/doi.org\/10.1600\/036364410790862678\">10.1600\/036364410790862678<\/a>, Accessed: 2025-07-30.<br \/>\n<span id=\"Attigala_2016\">Attigala L, Wysocki WP, Duvall MR and Clark LG (2016)<\/span> Phylogenetic estimation and morphological evolution of Arundinarieae (Bambusoideae:Poaceae) based on plastome phylogenomic analysis, Mol Phylog Evol, 101, 111-121, DOI: <a href=\"https:\/\/doi.org\/10.1016\/j.ympev.2016.05.008\">10.1016\/j.ympev.2016.05.008<\/a>, Accessed: 2025-07-30.<br \/>\n<span id=\"Takeda_1985\">\u6b66\u7530\u53cb\u56db\u90ce\u30fb\u8c37\u5ddd\u5b5d\u5f18\u30fb\u770c\u548c\u4e00\u30fb\u7bb1\u5c71\u664b (1985)<\/span> \u30a4\u30cd\u79d1C3, C4\u690d\u7269\u306e\u751f\u614b\u3068\u5730\u7406\u7684\u5206\u5e03\u306b\u95a2\u3059\u308b\u7814\u7a76, \u65e5\u4f5c\u7d00, 54(1), 54-64, DOI: <a href=\"https:\/\/doi.org\/10.1626\/jcs.54.54\">10.1626\/jcs.54.54<\/a>, Accessed: 2025-08-03.<br \/>\n<span id=\"Yoshimura_2015\">\u5409\u6751\u6cf0\u5e78 (2015)<\/span> \u56fd\u5185\u306b\u5206\u5e03\u3059\u308bC4\u690d\u7269\u306e\u30d5\u30ed\u30e9\u306e\u518d\u691c\u8a0e, \u65e5\u4f5c\u7d00, 84(4). 386\uff0d407, DOI: <a href=\"https:\/\/doi.org\/10.1626\/jcs.84.386\">10.1626\/jcs.84.386<\/a>, Accessed: 2025-08-04.<\/p>\n","protected":false},"excerpt":{"rendered":"<p>\u3000\u30a4\u30cd\u76ee(Poales)\u306f\uff12\u4e07\u7a2e\u4ee5\u4e0a\u304c\u8a18\u8f09\u3055\u308c\u3066\u6765\u305f\u5927\u898f\u6a21\u306a\u30b0\u30eb\u30fc\u30d7\u3067\u3001\u305d\u306e\u3046\u30611\u4e072000\u7a2e\u4ee5\u4e0a\u304c\u30a4\u30cd\u79d1\u306e\u6240\u5c5e\u3068\u306a\u3063\u3066\u3044\u307e\u3059(Wu et al.,2022)\u3002\u30a4\u30cd\u76ee\u5185\u306e\u30ab\u30c4\u30de\u30bf\u30bd\u30a6\u5468\u8fba\u306b\u8a8d\u3081\u3089\u308c\u305f3\u79d1\u304c\u7d71\u5408\u3055\u308c\u305f\u7d50\u679c\u3001 [&hellip;]<\/p>\n","protected":false},"author":1,"featured_media":0,"comment_status":"open","ping_status":"open","sticky":false,"template":"","format":"standard","meta":{"_jetpack_memberships_contains_paid_content":false,"footnotes":""},"categories":[13,7,2],"tags":[],"class_list":["post-52954","post","type-post","status-publish","format-standard","hentry","category-13","category-7","category-2"],"jetpack_featured_media_url":"","jetpack_sharing_enabled":true,"_links":{"self":[{"href":"https:\/\/ichihashi.me\/wp\/index.php?rest_route=\/wp\/v2\/posts\/52954","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/ichihashi.me\/wp\/index.php?rest_route=\/wp\/v2\/posts"}],"about":[{"href":"https:\/\/ichihashi.me\/wp\/index.php?rest_route=\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"https:\/\/ichihashi.me\/wp\/index.php?rest_route=\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"https:\/\/ichihashi.me\/wp\/index.php?rest_route=%2Fwp%2Fv2%2Fcomments&post=52954"}],"version-history":[{"count":2,"href":"https:\/\/ichihashi.me\/wp\/index.php?rest_route=\/wp\/v2\/posts\/52954\/revisions"}],"predecessor-version":[{"id":56734,"href":"https:\/\/ichihashi.me\/wp\/index.php?rest_route=\/wp\/v2\/posts\/52954\/revisions\/56734"}],"wp:attachment":[{"href":"https:\/\/ichihashi.me\/wp\/index.php?rest_route=%2Fwp%2Fv2%2Fmedia&parent=52954"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/ichihashi.me\/wp\/index.php?rest_route=%2Fwp%2Fv2%2Fcategories&post=52954"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/ichihashi.me\/wp\/index.php?rest_route=%2Fwp%2Fv2%2Ftags&post=52954"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}